Unia Europejska

foto Piotr Ślipiński

Progress in Studies on Myriapoda and Onychophora

Fragmenta faunistica, Supplement to Vol. 43
Edited by J. Wytwer and S. Golovatch Proceedings of the 11th International Congress of Myriapodology, 20-24 July 1999, Białowieża, Poland


The present volume puts on record the bulk, i.e. 33 papers by 52 authors from 19 countries, of the contributions delivered at the 11th International Congress of Myriapodology, held at Białowieża, Poland on 20-24 July 1999. This book is a milestone of recent progress achieved in research on Myriapoda and Onychophora, with the papers distributed between the following general topics:

  • Evolution, Phylogeny and Comparative Morphology (9 papers)
  • Anatomy, Biology and Physiology (10) 
  • Taxonomy and Collections (4) 
  • Geography, Faunistics and Ecology (10)

The book contains 14 introductory and 396 main text pages, 52 tables, 273 figures or graphs, and 2 indices.

The publication is available through the Library of the Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00-679 Warszawa, Poland, e-mail: Ten adres pocztowy jest chroniony przed spamowaniem. Aby go zobaczyć, konieczne jest włączenie w przeglądarce obsługi JavaScript.. One copy costs USD 50,-. Postage and handling charges are billed separately, the amount depending on the way of mailing. When ordering please specify the number of copies and the way of shipment. The corresponding invoice will be enclosed together with the shipment.

Fragmenta Faunistica

Editorial Office:

Museum and Institute of Zoology

Polish Academy of Sciences

Wilcza 64, 00-679 Warszawa, Poland


Editor in Chief:

Jolanta Wytwer

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Editorial Board:

Ewa Durska, Joanna Mazgajska, Wioletta Wawer

Advisory Board:

Józef Banaszak (Bydgoszcz, Poland)
Theo Blick (Hummeltal, Germany)
Sergey A. Belokobylskij (St. Petersburg, Russia)
Wiesław Bogdanowicz (Warszawa, Poland)
Thomas Bolger (Dublin, Ireland)
Jarosław Buszko (Toruń, Poland)
Adam Byk (Warszawa, Poland)
Wojciech Czechowski (Warszawa, Poland)
R. Henry L. Disney (Cambridge, UK)
Sergei Golovatch (Moscow, Russia)
Tomáš Lackner (Munich, Germany)
Abraham A. Mabelis (Wageningen, the Netherlands)
Wojciech Niedbała (Poznań, Poland)
Masahiro Ôhara (Sapporo, Japan)
Gilmar Perbiche-Neves (São Carlos, Brasil)
Václav Pižl (České Budéjovice, the Czech Republic)
Maria Sterzyńska (Warszawa, Poland)
Ryszard Szadziewski (Gdańsk, Poland)


All papers submitted to the Fragmenta Faunistica must reflect the author’s own research and analysis. It must not have been offered previously or simultaneously, to any other scientific magazine. Submission of a manuscript implies also that the publication has been approved by all co-authors, if any.

As of 2014, Fragmenta Faunistica applies the Creative Commons Attribution (CC BY) license to works. Under this license, authors retain ownership of the copyright for the content, but agree for anyone to download, reuse, reprint, modify, distribute and/or copy the content as long as the original authors and source are cited. No permission is required from the authors or the publishers in this case.

Manuscripts should be submitted to the Editor-in-Chief (Ten adres pocztowy jest chroniony przed spamowaniem. Aby go zobaczyć, konieczne jest włączenie w przeglądarce obsługi JavaScript.). A relevant letter from author(s) accompanying the manuscript is required where the contributions of all co-authors should be declared. The title page should include the names of the authors, the author’s address(es) in English, and email address of the corresponding author. The letter should also properly credit the contributions of co-authors and co-researchers.

Fragmenta Faunistica publishes in English. Papers should be concise and linguistically correct. Authors who are not native users of English are advised to have the manuscript linguistically reviewed before submitting it to the editor. Poorly written manuscripts will be returned without further review.

Manuscripts must conform to the following requirements:

Title should be concise and relevant to its contents. Abstract should explain the idea of the research and summarize the results, not to exceed 200 words and should not contain any references. Key words numbering 3 to 7 should not be repeated in the title. Text should have a standard layout (introduction, material and methods, results, discussion, possible conclusions, acknowledgements, references) and a brief but comprehensive summary, which will be translated into Polish. Names of genera and species must be italicised. The author’s name must be added to each species (genus) names first appearing in the text or be incorporated into one of the tables in the paper. Tables should be as simple as possible, should contain captions and be numbered (cited as ‘Table 1’ or ‘Tables 1 & 2’). Illustrations should be submitted separately as graphic files (>300 dpi) and be numbered as successive figures (cited as ‘Fig. 1’ or ‘Figs 1 & 2’). Numerical data should be attached to plots. All captions should be enclosed to the manuscript. Picture credits should be given as ‘Photo by J. Bean’. Colour illustrations may be accepted by the editor. Dates should be given as follows, e.g. 1−2 Feb 2014. Polish summary is required. Non-Polish authors should prepare an English summary.

Citations in the text should be written as follows: Thayer (1992), Gamarra & Outerelo (2008) or (Thayer 1992, Assing 2009), where the order is chronological. The authors’ names should be separated by coma or by semicolon if citation includes more than one work of one author, i.e.: (Disney 1989, 1990, 1997; Liu 2001; Disney & Bänziger 2009). Publications with more than two authors should be cited in the text with the first author and the annotation ‘et al.’ i.e. Mossakowski et al. (2003). In exceptional cases, if data have no reference, it should be cited as: ‘D. Spiller, pers. comm.’ or ‘pers. obs.’ or ‘unpubl. data’.

References should be listed in alphabetical order; book and journal titles must be given in full. If available, please always include DOIs as full DOI links in reference list. Non-Latin names of journals should be given according the BIOSIS Base Source. The title of the publication should be given in English according to their edition details and if it is impossible - transliterate; in the latter case, the author provides his own translation of the title, placing it immediately after it in square brackets. The indication of the original language should be put in square brackets at the end of the referenced item. A few examples of references you find here.

Principles of reviewing procedure. The Editor-in-Chief may optionally make a quick assessment on the suitability of the paper for Fragmenta Faunistica. Papers can be rejected or returned for revision to the authors at this stage without full formal review. Each manuscript qualified for further assessment is reviewed by at least two independent reviewers from outside the institutions in which the author is affiliated. One reviewer must be of another nationality than the author. Reviewers declare no conflict of interest with author. Review is provided as a filled form with a clear statement on acceptance or rejection. In the case of two very different opinions, the Editor-in-Chief may obtained a review from the third reviewer. Manuscripts can only be accepted after two positive reviews. The Author receives anonymous reviews. The final version of the manuscript should include both the comments of reviewers and editors. Rejected papers are archived for five years in the Editorial Board.

Subscriptions and orders should be addressed to the Museum & Institute of Zoology, PAS, Library, Wilcza 64, 00-679 Warszawa. Annual subscription (2 issues) institutional:  EURO 60, personal: EURO 40. Orders paid on the account on: Museum & Institute of Zoology, PAS, Wilcza 64, 00679 Warszawa,

BGK IBAN PL 24 1130 1017 0020 1465 5520 0001 (PLN)

BGK IBAN PL 93 1130 1017 0020 1465 5520 0020 (EUR)

BGK IBAN PL 23 1130 1017 0020 1465 (USD)






An International Journal of Faunology


ISSN 0015-9301

eISSN 2391-5846

Fragmenta Faunistica is an international, semiannual journal published by the Museum and Institute of Zoology of the Polish Academy of Sciences. The journal, first published in 1930, appeared under the title Fragmenta Faunistica Musei Zoologici Polonici until 1953. Now, it is a peer reviewed, open accessjournal, publishing the papers devoted to fauna – its differentiation, distribution and transformations. Faunology, which quantitatively examines the dynamics and structure of faunistic complexes is the main object. The journal publishes the results of original studies, review articles and syntheses dealing with faunology and with related sciences such as zoogeography, zoocoenology or taxonomy.

Fragmenta Faunistica is indexed in: AGRO, Biological Abstracts, BIOSIS Preview/BIOSIS, Google Scholar, IC Journals Master List (ICV 2020 = 93.71), Worldcat, Zoological Record. Ministry of Education and Science in Poland (20 points).

Fragmenta Faunistica is available from the platform of the Digital Repository of Scientific Institutes:https://rcin.org.pl/